cryogenic (thermokarst) lakes
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Lakes formed when permafrost thaws and ground collapses.
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cryogenic (thermokarst) lakes
Lakes formed when permafrost thaws and ground collapses.
volcanic crater lake / caldera lake
Lakes forming in volcano craters or collapsed volcanic cones.
fault-derived basins (Graben)
Lakes formed along tectonic faulting, creating deep, steep-walled basins.
littoral zone
Shallow nearshore region where light reaches the bottom, allowing growth of rooted macrophytes.
pelagic zone
Open-water region away from shore, extending down to the compensation depth.
profundal zone
Deep, aphotic bottom region below the compensation depth; inhabited by benthic organisms.
benthos
Organisms living on or in lake sediments (worms, insect larvae, mollusks).
plankton
Drifting organisms with limited ability to swim (phytoplankton + zooplankton).
euphotic zone
Upper layer where photosynthesis > respiration. Extends from surface to compensation depth.
compensation depth
Depth at which photosynthetic oxygen production exactly equals respiration.
redox potential (Eh)
Measure of the tendency of the environment to accept or donate electrons.
ammonification
Conversion of organic N to NH4+ by microbes.
nitrification
Conversion of NH4+ → NO2- → NO3- under aerobic conditions.
denitrification
Anaerobic reduction of NO3- to N2 gas.
N-fixation
Conversion of N2 gas to NH4+ by cyanobacteria using nitrogenase.
dissolved organic nitrogen (DON)
Organic N compounds dissolved in water, including urea and amino acids.
dissolved organic phosphorus (DOP)
Organic P compounds in the water column.
luxury uptake
Algae store excess P when available and use it later during deficiency.
gross primary production (GPP)
Total photosynthesis unchecked by respiration.
net primary production (NPP)
GPP minus phytoplankton respiration.
standing crop
Total amount of living organisms in a specific area at one time.
P/B ratio
Production-to-biomass ratio; indicates turnover rate.
O2-change method
Measures primary production via changes in dissolved oxygen in light and dark bottles.
flux
F = D A C / z (mass transfer rate) describing diffusion-driven exchange.
moraine lakes
Lakes dammed by glacial moraines (debris left by glaciers).
piston velocity
Gas transfer velocity describing how quickly gases exchange across the surface film.
catchment/watershed
Land area draining water into a lake or river.
baseflow
Portion of streamflow supplied by slow groundwater discharge.
stormflow
Runoff entering streams during and shortly after rainfall events.
water yield
Amount of water produced by a landscape (typically measured as runoff).
endorheic basin
Drainage basin that does not drain to the ocean; promotes saline lakes.
exorheic basin
Drainage basin where rivers ultimately reach the ocean.
potential evapotranspiration (PET)
Energy-controlled maximum evaporation possible from landscape surfaces.
solution lakes
Formed when soluble rock (e.g., limestone) dissolves, creating depressions.
oxbow lake
U-shaped lake formed when a meandering river bend is cut off.
meander scroll lake
Small lakes formed on floodplains due to meandering river migration.
plunge-pool lake
Lake formed by erosional scouring beneath waterfalls.
glacial rock basins
Basins carved by glaciers, forming various lake types.
ice-scour lakes
Formed by valley glaciers gouging bedrock.
cirque lakes
Formed at the head of glacial valleys in amphitheater-like hollows.
fjord lakes
Glacial valleys flooded by seawater and later uplifted.
nutrient
A substance that provides food, allowing organisms to grow and survive.
aphotic zone
The layer of water that extends from the photic zone to the bottom of the lake.
net ecosystem production
The difference between gross primary production and cellular respiration; a measure of how much nutrients, especially carbon, are in the system.
autochthonous production
Matter that is produced from within the ecosystem.
meromictic lake
Lakes that are permanently stratified with a monimolimnion layer (a low-oxygen layer, usually near the bottom).
nekton
An independent community of organisms able to move independently by swimming.
kettle lakes
Lakes formed from melting blocks of ice trapped within a ground moraine.
shear
Breakage of a material, generally due to compressive stress; ductile materials shift, brittle materials form fault zones.
heterocyst
Thick-walled cells in algae that create low-oxygen zones for nitrogen fixation; lack photosystem II and have adaptations to limit O2.
"drift organisms"
Organisms that are passively moved around by water currents (microscopic organisms and some invertebrates).
extinction coefficient (k)
Rate at which light intensity decreases with depth due to absorption and scattering.
albedo
Fraction of incoming light reflected off the water surface.
net radiation (QN)
Balance of incoming and outgoing shortwave and longwave radiation at the lake surface.
sensible heat transfer (QH)
Heat exchange between lake and atmosphere due to temperature differences.
latent heat flux (QE)
Heat used for evaporation from the lake surface.
stagnant-film model
Model describing gas exchange across the thin, stagnant layer at the air-water interface.
14C uptake method
Radiolabelling technique used to measure photosynthesis (measures carbon assimilation using radioactive carbonate).
Henry's Law
Solubility of a gas is proportional to its partial pressure.
eddy diffusion / turbulent diffusion
Mixing generated by turbulent eddies, especially strong horizontally.
fetch
Distance wind blows over water; controls wave formation.
Langmuir circulation
Rotating water at lake surface; wind-driven spiral cells creating surface streaks of foam and alternating upwelling/downwelling zones.
surface seiche
Standing wave caused by wind piling water on one side of the lake.
internal seiche
Oscillating wave of the thermocline in stratified lakes.
laminar flow
Water moves in smooth, parallel layers with little mixing; occurs at low Reynolds number.
turbulent flow
Chaotic, eddy-filled motion with strong mixing; occurs at high Reynolds number.
Reynolds number (Re)
Predicts laminar vs turbulent flow: Re = U d / ν.
Richardson number (Ri)
Predicts whether stratification suppresses mixing.
advection
Bulk horizontal transport of water and its contents.
epilimnion
Warm, well-mixed upper layer.
hypolimnion
Cold, dense, largely isolated deep layer.
thermocline / metalimnion
Layer of rapid temperature change separating epilimnion and hypolimnion.
holomictic lake
Lake that mixes completely at least once a year.
amictic lake
Never mixes (usually permanently ice-covered in polar regions).
dimictic lake
Mixes twice per year (spring and fall).
monomictic lake
Mixes once per year.
polymictic lake
Mixes many times per year (shallow tropical lakes).
oligomictic lake
Mixes irregularly or rarely due to warmth in tropical settings.
planktivory
Feeding on zooplankton.
herbivory (fish)
Feeding on periphyton or filamentous algae.
detritivory
Feeding on organic detritus.
keystone predator
Predator exerting disproportionately large control on community structure.
trophic cascade
Top-down effects where predator abundance affects herbivores and primary producers.
multivoltine species
Having multiple generations per year (rotifers, cladocerans).
univoltine species
Only one brood generation per year (some copepods).
filter feeding
Capturing suspended food particles using appendages; influenced by Reynolds constraints.
cyclomorphosis
Seasonal morphological changes in zooplankton (e.g., helmets, spines).
diel vertical migration
Daily movement up at night and down during the day to balance feeding vs predation risk.
solar bottleneck hypothesis
High UV exposure near surface constrains vertical distribution of plankton.
microbial loop
Pathway where DOC → bacteria → protozoa → zooplankton, recycling energy and nutrients.
chromophoric DOC (CDOM)
DOC that absorbs UV/visible light, affecting light penetration and photochemistry.
photochemical breakdown of DOC
UV-driven reactions converting CDOM into more labile carbon for bacteria.
holoplanktonic diatoms
Diatoms that remain suspended; they bloom in spring.
meroplanktonic diatoms
Diatoms with a benthic resting phase; resuspended during fall/winter mixing.
critical depth
Depth where integrated photosynthesis equals integrated respiration.
critical turbulence
Level of turbulence needed to keep cells in the euphotic zone.
Monod equation
Describes growth vs nutrient concentration: μ = μmax S / (Ks + S); Ks is the half-saturation constant.
Ks (half-saturation constant)
Nutrient concentration at which growth is half maximal.
alkaline phosphatase
Enzyme that breaks down organic P to inorganic phosphate; produced under P limitation to liberate PO4³⁻ from organic compounds.
internal loading
Release of P from anoxic sediments back into the water column.